I was doing some scrys when I started noticing that Guardian Males have a lot of uncolored or misaligned pixels between the primary/secondary colors on a lot of genes. Most of it seems to be a base layer issue (like with the tail and back spines) though there are some that seem gene-unique (or can be found easier).
Basic/Basic for comparison. There is a single uncolored primary pixel in the middle of where the wing connects to the rib ridge.
Secondary color bleeding/primary color receding issues:
At the very top of the thigh where the lineart curves into the spine and along the top tail lineart
Along the "arm" of the front wing
Where the beard meets the chin
Back wing shoulder (or it at least looks "messy")
The two back spines that overlap the back wing membrane (those light pixels will go dark if you switch to a dark color like Phthalo)
Piebald - the back spines closest to the wings have a little white edge on them.
Blaze - same issues as Basic + lineart halo on the front shoulder nearest the back spines
Facet - same issues as Basic + stronger pixel halo on back spines now includes a 3rd spine (the one in front of the back wing's arm)
Fissure - additional bleed around the rib-attachment area
Noxtide - additional bleed around the rib-attachment area and missing color from around the 3 back spines (could be an issue with base primary layer, idk)
Paint - white pixels along back between 2 back spines (but is not in a location colored white by the paint gene)
Saddle - where I first noticed the issue - pixel issues are across the entire wing where it meets the body (tail, lower back, rib ridge, front wing arm (seen better on grapefruit), both wing shoulders)
Sarcophagus - likely has same base layer issues as seen by the origin color bleeding through in select areas (between back spines, front wing arm)
Toxin - another example where the back spine pixel issue is more visible
Weaver - pixel halo on back wing shoulder is very prominent